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T4 RNA Ligase 2, also known as T4 Rnl2 (gp24.1), has both intermolecular and intramolecular RNA strand joining activity

Unlike T4 RNA Ligase 1, T4 RNA Ligase 2 is much more active joining nicks on double stranded RNA than on joining the ends of single stranded RNA. The enzyme requires an adjacent 5′ phosphate and 3′ OH for ligation. The enzyme can also ligate the 3′ OH of RNA to the 5′ phosphate of DNA in a double stranded structure.